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Fungal fossils do not become common and uncontroversial until the early Devonian (416–359.2 Ma), when they are abundant in the Rhynie chert, mostly as Zygomycota and Chytridiomycota.

Fungal fossils are difficult to distinguish from those of other microbes, and are most easily identified when they resemble extant fungi.

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Since the fungi form a sister group to the animals, the two lineages must have diverged before the first animal lineages, which are known from fossils as early as the Ediacaran.

In contrast to plants and animals, the early fossil record of the fungi is meager.

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